Carcasses of hunted wolves in Volgograd Oblast, Russia
Theodore Roosevelt said wolves are difficult to hunt because of their elusiveness, sharp senses, high endurance, and ability to quickly incapacitate and kill hunting dogs.[202] Historic methods included killing of spring-born litters in their dens, coursing with dogs (usually combinations of sighthounds, Bloodhounds and Fox Terriers), poisoning with strychnine, and trapping.[203][204]
A popular method of wolf hunting in Russia involves trapping a pack within a small area by encircling it with fladry poles carrying a human scent. This method relies heavily on the wolf's fear of human scents, though it can lose its effectiveness when wolves become accustomed to the odor. Some hunters can lure wolves by imitating their calls. In Kazakhstan and Mongolia, wolves are traditionally hunted using eagles and large falcons, though this practice is declining, as experienced falconers are becoming few in number. Shooting wolves from aircraft is highly effective, due to increased visibility and direct lines of fire.[204] Several types of dog, including the Borzoi and Kyrgyz Tajgan, have been specifically bred for wolf hunting.[192]
Wolves as pets and working animals[edit]
Wolves and wolf-dog hybrids are sometimes kept as exotic pets, wolves do not show the same tractability as dogs in living alongside humans, being generally less responsive to human commands and more likely to act aggressively. Humans are more likely to be fatally mauled by a pet wolf or wolf-dog hybrid than by a dog.[205]
![]()
^ Populations of Bhutan, India, Nepal, and Pakistan are included in Appendix I. Excludes domesticated form and dingo, which are referenced as Canus lupus familiaris and Canus lupus dingo.
^ Domestic and feral dogs are included in the phylogenetic but not colloquial definition of 'wolf', and are thus not in the scope of this article.
^ a b c d e Boitani, L.; Phillips, M. & Jhala, Y. (2023) [amended version of 2018 assessment]. "Canis lupus". IUCN Red List of Threatened Species. 2023 e.T3746A247624660. doi:10.2305/IUCN.UK.2023-1.RLTS.T3746A247624660.en. Retrieved June 2, 2024.
^ "Appendices | CITES". cites.org. Retrieved January 14, 2022.
^ a b c Linnæus, Carl (1758). "Canis Lupus". Systema naturæ per regna tria naturæ, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I (in Latin) (10 ed.). Holmiæ (Stockholm): Laurentius Salvius. pp. 39–40.
^ Harper, Douglas. "wolf". Online Etymology Dictionary.
^ A. Lehrman (1987). "Anatolian Cognates of the PIE Word for 'Wolf'". Die Sprache. 33: 13–18.
^ Young, Stanley P.; Goldman, Edward A. (1944). The Wolves of North America, Part I. New York, Dover Publications, Inc. p. 390.
^ Marvin 2012, pp. 74–75.
^ Koepfli, Klaus-Peter; Pollinger, John; Godinho, Raquel; Robinson, Jacqueline; Lea, Amanda; Hendricks, Sarah; Schweizer, Rena M.; Thalmann, Olaf; Silva, Pedro; Fan, Zhenxin; Yurchenko, Andrey A.; Dobrynin, Pavel; Makunin, Alexey; Cahill, James A.; Shapiro, Beth; Álvares, Francisco; Brito, José C.; Geffen, Eli; Leonard, Jennifer A.; Helgen, Kristofer M.; Johnson, Warren E.; o'Brien, Stephen J.; Van Valkenburgh, Blaire; Wayne, Robert K. (2015). "Genome-wide Evidence Reveals that African and Eurasian Golden Jackals Are Distinct Species". Current Biology. 25 (#16): 2158–65. Bibcode:2015CBio...25.2158K. doi:10.1016/j.cub.2015.06.060. PMID 26234211.
^ Harper, Douglas. "canine". Online Etymology Dictionary.
^ Clutton-Brock, Juliet (1995). "2-Origins of the dog". In Serpell, James (ed.). The Domestic Dog: Its Evolution, Behaviour and Interactions with People. Cambridge University Press. pp. 7–20. ISBN 0-521-41529-2.
^ Wozencraft, W. C. (2005). "Order Carnivora". In Wilson, D. E.; Reeder, D. M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. pp. 575–577. ISBN 978-0-8018-8221-0. OCLC 62265494.
^ Larson, G.; Bradley, D. G. (2014). "How Much Is That in Dog Years? The Advent of Canine Population Genomics". PLOS Genetics. 10 (1) e1004093. doi:10.1371/journal.pgen.1004093. PMC 3894154. PMID 24453989.
^ Alvares, Francisco; Bogdanowicz, Wieslaw; Campbell, Liz A.D.; Godinho, Rachel; Hatlauf, Jennifer; Jhala, Yadvendradev V.; Kitchener, Andrew C.; Koepfli, Klaus-Peter; Krofel, Miha; Moehlman, Patricia D.; Senn, Helen; Sillero-Zubiri, Claudio; Viranta, Suvi; Werhahn, Geraldine (2019). "Old World Canis spp. with taxonomic ambiguity: Workshop conclusions and recommendations. CIBIO. Vairão, Portugal, 28–30 May 2019" (PDF). IUCN/SSC Canid Specialist Group. Retrieved March 6, 2020.
^ Mech & Boitani 2003, pp. 239–245.
^ a b Iurino, Dawid A.; Mecozzi, Beniamino; Iannucci, Alessio; Moscarella, Alfio; Strani, Flavia; Bona, Fabio; Gaeta, Mario; Sardella, Raffaele (February 25, 2022). "A Middle Pleistocene wolf from central Italy provides insights on the first occurrence of Canis lupus in Europe". Scientific Reports. 12 (1): 2882. Bibcode:2022NatSR..12.2882I. doi:10.1038/s41598-022-06812-5. ISSN 2045-2322. PMC 8881584. PMID 35217686.
^ Tedford, Richard H.; Wang, Xiaoming; Taylor, Beryl E. (2009). "Phylogenetic Systematics of the North American Fossil Caninae (Carnivora: Canidae)". Bulletin of the American Museum of Natural History. 325: 1–218. doi:10.1206/574.1. hdl:2246/5999. S2CID 83594819.
^ a b c Thalmann, Olaf; Perri, Angela R. (2018). "Paleogenomic Inferences of Dog Domestication". In Lindqvist, C.; Rajora, O. (eds.). Paleogenomics. Population Genomics. Springer, Cham. pp. 273–306. doi:10.1007/13836_2018_27. ISBN 978-3-030-04752-8.
^ Freedman, Adam H.; Gronau, Ilan; Schweizer, Rena M.; Ortega-Del Vecchyo, Diego; Han, Eunjung; et al. (2014). "Genome Sequencing Highlights the Dynamic Early History of Dogs". PLOS Genetics. 10 (1). e1004016. doi:10.1371/journal.pgen.1004016. PMC 3894170. PMID 24453982.
^ a b c Fan, Zhenxin; Silva, Pedro; Gronau, Ilan; Wang, Shuoguo; Armero, Aitor Serres; Schweizer, Rena M.; Ramirez, Oscar; Pollinger, John; Galaverni, Marco; Ortega Del-Vecchyo, Diego; Du, Lianming; Zhang, Wenping; Zhang, Zhihe; Xing, Jinchuan; Vilà, Carles; Marques-Bonet, Tomas; Godinho, Raquel; Yue, Bisong; Wayne, Robert K. (2016). "Worldwide patterns of genomic variation and admixture in gray wolves". Genome Research. 26 (2): 163–173. doi:10.1101/gr.197517.115. PMC 4728369. PMID 26680994.
^ Hennelly, Lauren M.; Habib, Bilal; Modi, Shrushti; Rueness, Eli K.; Gaubert, Philippe; Sacks, Benjamin N. (2021). "Ancient divergence of Indian and Tibetan wolves revealed by recombination-aware phylogenomics". Molecular Ecology. 30 (24): 6687–6700. Bibcode:2021MolEc..30.6687H. doi:10.1111/mec.16127. PMID 34398980. S2CID 237147842.
![]()
^ a b Bergström, Anders; Stanton, David W. G.; Taron, Ulrike H.; Frantz, Laurent; Sinding, Mikkel-Holger S.; Ersmark, Erik; Pfrengle, Saskia; Cassatt-Johnstone, Molly; Lebrasseur, Ophélie; Girdland-Flink, Linus; Fernandes, Daniel M.; Ollivier, Morgane; Speidel, Leo; Gopalakrishnan, Shyam; Westbury, Michael V. (July 14, 2022). "Grey wolf genomic history reveals a dual ancestry of dogs". Nature. 607 (7918): 313–320. Bibcode:2022Natur.607..313B. doi:10.1038/s41586-022-04824-9. ISSN 0028-0836. PMC 9279150. PMID 35768506.
^ a b Loog, Liisa; Thalmann, Olaf; Sinding, Mikkel-Holger S.; Schuenemann, Verena J.; Perri, Angela; Germonpré, Mietje; Bocherens, Herve; Witt, Kelsey E.; Samaniego Castruita, Jose A.; Velasco, Marcela S.; Lundstrøm, Inge K. C.; Wales, Nathan; Sonet, Gontran; Frantz, Laurent; Schroeder, Hannes (May 2020). "Ancient DNA suggests modern wolves trace their origin to a Late Pleistocene expansion from Beringia". Molecular Ecology. 29 (9): 1596–1610. Bibcode:2020MolEc..29.1596L. doi:10.1111/mec.15329. ISSN 0962-1083. PMC 7317801. PMID 31840921.
^ Freedman, Adam H; Wayne, Robert K (2017). "Deciphering the Origin of Dogs: From Fossils to Genomes". Annual Review of Animal Biosciences. 5: 281–307. doi:10.1146/annurev-animal-022114-110937. PMID 27912242. S2CID 26721918.
^ a b Gopalakrishnan, Shyam; Sinding, Mikkel-Holger S.; Ramos-Madrigal, Jazmín; Niemann, Jonas; Samaniego Castruita, Jose A.; Vieira, Filipe G.; Carøe, Christian; Montero, Marc de Manuel; Kuderna, Lukas; Serres, Aitor; González-Basallote, Víctor Manuel; Liu, Yan-Hu; Wang, Guo-Dong; Marques-Bonet, Tomas; Mirarab, Siavash; Fernandes, Carlos; Gaubert, Philippe; Koepfli, Klaus-Peter; Budd, Jane; Rueness, Eli Knispel; Heide-Jørgensen, Mads Peter; Petersen, Bent; Sicheritz-Ponten, Thomas; Bachmann, Lutz; Wiig, Øystein; Hansen, Anders J.; Gilbert, M. Thomas P. (2018). "Interspecific Gene Flow Shaped the Evolution of the Genus Canis". Current Biology. 28 (21): 3441–3449.e5. Bibcode:2018CBio...28E3441G. doi:10.1016/j.cub.2018.08.041. PMC 6224481. PMID 30344120.
^ a b Sinding, Mikkel-Holger S.; Gopalakrishan, Shyam; Vieira, Filipe G.; Samaniego Castruita, Jose A.; Raundrup, Katrine; Heide Jørgensen, Mads Peter; Meldgaard, Morten; Petersen, Bent; Sicheritz-Ponten, Thomas; Mikkelsen, Johan Brus; Marquard-Petersen, Ulf; Dietz, Rune; Sonne, Christian; Dalén, Love; Bachmann, Lutz; Wiig, Øystein; Hansen, Anders J.; Gilbert, M. Thomas P. (2018). "Population genomics of grey wolves and wolf-like canids in North America". PLOS Genetics. 14 (11) e1007745. doi:10.1371/journal.pgen.1007745. PMC 6231604. PMID 30419012.
^ Wang, Guo-Dong; Zhang, Ming; Wang, Xuan; Yang, Melinda A.; Cao, Peng; Liu, Feng; Lu, Heng; Feng, Xiaotian; Skoglund, Pontus; Wang, Lu; Fu, Qiaomei; Zhang, Ya-Ping (2019). "Genomic Approaches Reveal an Endemic Subpopulation of Gray Wolves in Southern China". iScience. 20: 110–118. Bibcode:2019iSci...20..110W. doi:10.1016/j.isci.2019.09.008. PMC 6817678. PMID 31563851.
^ Iacolina, Laura; Scandura, Massimo; Gazzola, Andrea; Cappai, Nadia; Capitani, Claudia; Mattioli, Luca; Vercillo, Francesca; Apollonio, Marco (2010). "Y-chromosome microsatellite variation in Italian wolves: A contribution to the study of wolf-dog hybridization patterns". Mammalian Biology—Zeitschrift für Säugetierkunde. 75 (4): 341–347. Bibcode:2010MamBi..75..341I. doi:10.1016/j.mambio.2010.02.004.
^ a b Kopaliani, N.; Shakarashvili, M.; Gurielidze, Z.; Qurkhuli, T.; Tarkhnishvili, D. (2014). "Gene Flow between Wolf and Shepherd Dog Populations in Georgia (Caucasus)". Journal of Heredity. 105 (3): 345–53. doi:10.1093/jhered/esu014. PMID 24622972.
^ Moura, A. E.; Tsingarska, E.; Dąbrowski, M. J.; Czarnomska, S. D.; Jędrzejewska, B. A.; Pilot, M. G. (2013). "Unregulated hunting and genetic recovery from a severe population decline: The cautionary case of Bulgarian wolves". Conservation Genetics. 15 (2): 405–417. doi:10.1007/s10592-013-0547-y.
^ Bergström, Anders; Frantz, Laurent; Schmidt, Ryan; Ersmark, Erik; Lebrasseur, Ophelie; Girdland-Flink, Linus; Lin, Audrey T.; Storå, Jan; Sjögren, Karl-Göran; Anthony, David; Antipina, Ekaterina; Amiri, Sarieh; Bar-Oz, Guy; Bazaliiskii, Vladimir I.; Bulatović, Jelena; Brown, Dorcas; Carmagnini, Alberto; Davy, Tom; Fedorov, Sergey; Fiore, Ivana; Fulton, Deirdre; Germonpré, Mietje; Haile, James; Irving-Pease, Evan K.; Jamieson, Alexandra; Janssens, Luc; Kirillova, Irina; Horwitz, Liora Kolska; Kuzmanovic-Cvetković, Julka; Kuzmin, Yaroslav; Losey, Robert J.; Dizdar, Daria Ložnjak; Mashkour, Marjan; Novak, Mario; Onar, Vedat; Orton, David; Pasaric, Maja; Radivojevic, Miljana; Rajkovic, Dragana; Roberts, Benjamin; Ryan, Hannah; Sablin, Mikhail; Shidlovskiy, Fedor; Stojanovic, Ivana; Tagliacozzo, Antonio; Trantalidou, Katerina; Ullén, Inga; Villaluenga, Aritza; Wapnish, Paula; Dobney, Keith; Götherström, Anders; Linderholm, Anna; Dalén, Love; Pinhasi, Ron; Larson, Greger; Skoglund, Pontus (2020). "Origins and genetic legacy of prehistoric dogs". Science. 370 (6516): 557–564. doi:10.1126/science.aba9572. PMC 7116352. PMID 33122379. S2CID 225956269.
^ a b Mech, L. David (1974). "Canis lupus". Mammalian Species (37): 1–6. doi:10.2307/3503924. JSTOR 3503924. Archived from the original on July 31, 2019. Retrieved July 30, 2019.
^ Heptner & Naumov 1998, pp. 129–132.
^ a b Heptner & Naumov 1998, p. 166.
^ Mech 1981, p. 13.
^ Tomiya, Susumu; Meachen, Julie A. (January 17, 2018). "Postcranial diversity and recent ecomorphic impoverishment of North American gray wolves". Biology Letters. 14 (1) 20170613. doi:10.1098/rsbl.2017.0613. ISSN 1744-9561. PMC 5803591. PMID 29343558.
^ a b c d e f g h i j k l m n o p q Heptner & Naumov 1998, pp. 164–270.
^ Mech 1981, p. 14.
^ Therrien, F. O. (2005). "Mandibular force profiles of extant carnivorans and implications for the feeding behaviour of extinct predators". Journal of Zoology. 267 (3): 249–270. doi:10.1017/S0952836905007430.
^ Mech & Boitani 2003, p. 112.
^ a b c d e f g h Paquet, P.; Carbyn, L. W. (2003). "Ch23: Gray wolf Canis lupus and allies". In Feldhamer, G. A.; Thompson, B. C.; Chapman, J. A. (eds.). Wild Mammals of North America: Biology, Management, and Conservation (2 ed.). Johns Hopkins University Press. pp. 482–510. ISBN 0-8018-7416-5.[permanent dead link]
^ a b Lopez 1978, p. 23.
^ a b Heptner & Naumov 1998, p. 174.
![]()
^ a b Miklosi, A. (2015). "Ch. 5.5.2—Wolves". Dog Behaviour, Evolution, and Cognition (2 ed.). Oxford University Press. pp. 110–112. ISBN 978-0-19-104572-1.
^ Macdonald, D. W.; Norris, S. (2001). Encyclopedia of Mammals. Oxford University Press. p. 45. ISBN 978-0-7607-1969-5.
^ a b c Lopez 1978, p. 19.
^ Lopez 1978, p. 18.
^ Lopez 1978, pp. 19–20.
^ a b Gipson, Philip S.; Bangs, Edward E.; Bailey, Theodore N.; Boyd, Diane K.; Cluff, H. Dean; Smith, Douglas W.; Jiminez, Michael D. (2002). "Color Patterns among Wolves in Western North America". Wildlife Society Bulletin. 30 (3): 821–830. JSTOR 3784236.
^ Heptner & Naumov 1998, pp. 168–169.
^ Heptner & Naumov 1998, p. 168.
^ Lopez 1978, p. 22.
^ Anderson, T. M.; Vonholdt, B. M.; Candille, S. I.; Musiani, M.; Greco, C.; Stahler, D. R.; Smith, D. W.; Padhukasahasram, B.; Randi, E.; Leonard, J. A.; Bustamante, C. D.; Ostrander, E. A.; Tang, H.; Wayne, R. K.; Barsh, G. S. (2009). "Molecular and Evolutionary History of Melanism in North American Gray Wolves". Science. 323 (5919): 1339–1343. Bibcode:2009Sci...323.1339A. doi:10.1126/science.1165448. PMC 2903542. PMID 19197024.
^ Hedrick, P. W. (2009). "Wolf of a different colour". Heredity. 103 (6): 435–436. Bibcode:2009Hered.103..435H. doi:10.1038/hdy.2009.77. PMID 19603061. S2CID 5228987.
^ Earle, M (1987). "A flexible body mass in social carnivores". American Naturalist. 129 (5): 755–760. Bibcode:1987ANat..129..755E. doi:10.1086/284670. S2CID 85236511.
^ Sorkin, Boris (2008). "A biomechanical constraint on body mass in terrestrial mammalian predators". Lethaia. 41 (4): 333–347. Bibcode:2008Letha..41..333S. doi:10.1111/j.1502-3931.2007.00091.x.
^ Mech, L. David (1966). The Wolves of Isle Royale. Fauna Series 7. Fauna of the National Parks of the United States. pp. 75–76. ISBN 978-1-4102-0249-9.
^ a b Newsome, Thomas M.; Boitani, Luigi; Chapron, Guillaume; Ciucci, Paolo; Dickman, Christopher R.; Dellinger, Justin A.; López-Bao, José V.; Peterson, Rolf O.; Shores, Carolyn R.; Wirsing, Aaron J.; Ripple, William J. (2016). "Food habits of the world's grey wolves". Mammal Review. 46 (4): 255–269. Bibcode:2016MamRv..46..255N. doi:10.1111/mam.12067. hdl:10536/DRO/DU:30085823. S2CID 31174275.
^ a b Mech & Boitani 2003, p. 107.
^ Mech & Boitani 2003, pp. 109–110.
^ Landry, Zoe; Kim, Sora; Trayler, Robin B.; Gilbert, Marisa; Zazula, Grant; Southon, John; Fraser, Danielle (June 1, 2021). "Dietary reconstruction and evidence of prey shifting in Pleistocene and recent gray wolves (Canis lupus) from Yukon Territory". Palaeogeography, Palaeoclimatology, Palaeoecology. 571 110368. Bibcode:2021PPP...57110368L. doi:10.1016/j.palaeo.2021.110368. ISSN 0031-0182. Retrieved April 23, 2024 – via Elsevier Science Direct.
^ Mech 1981, p. 172.
^ a b Mech & Boitani 2003, p. 201.
^ a b c d Heptner & Naumov 1998, pp. 213–231.
^ Barton, Brandon T.; Hill, JoVonn G.; Wolff, Carter L.; Newsome, Thomas M.; Ripple, William J.; Lashley, Marcus A. (September 18, 2019). "Grasshopper consumption by grey wolves and implications for ecosystems". Ecology. 101 (2) e02892. doi:10.1002/ecy.2892. ISSN 0012-9658. PMID 31531974. Retrieved April 20, 2025 – via Wiley Online Library.
![]()
^ Gable, T. D.; Windels, S. K.; Homkes, A. T. (2018). "Do wolves hunt freshwater fish in spring as a food source?". Mammalian Biology. 91: 30–33. Bibcode:2018MamBi..91...30G. doi:10.1016/j.mambio.2018.03.007. S2CID 91073874.
^ Woodford, Riley (November 2004). "Alaska's Salmon-Eating Wolves". Wildlifenews.alaska.gov. Retrieved July 25, 2019.
^ McAllister, I. (2007). The Last Wild Wolves: Ghosts of the Rain Forest. University of California Press. p. 144. ISBN 978-0-520-25473-2.
^ Fuller, T. K. (2019). "Ch3-What wolves eat". Wolves: Spirit of the Wild. Chartwell Crestline. p. 53. ISBN 978-0-7858-3738-1.
^ Mech & Boitani 2003, p. 109.
^ Mech 1981, p. 180.
^ Klein, D. R. (1995). "The introduction, increase, and demise of wolves on Coronation Island, Alaska". In Carbyn, L. N.; Fritts, S. H.; Seip, D. R. (eds.). Ecology and conservation of wolves in a changing world. Canadian Circumpolar Institute, Occasional Publication No. 35. pp. 275–280.
^ a b Mech & Boitani 2003, pp. 266–268.
^ Merrit, Dixon (January 7, 1921). "World's Greatest Animal Dead" (PDF). US Department of Agriculture Division of Publications. p. 2. Archived (PDF) from the original on July 24, 2019. Retrieved July 26, 2019.
^ Giannatos, G. (April 2004). "Conservation Action Plan for the golden jackal Canis aureus L. in Greece" (PDF). World Wildlife Fund Greece. pp. 1–47. Archived (PDF) from the original on December 9, 2017. Retrieved October 29, 2019.
^ Mech & Boitani 2003, p. 269.
^ Mech & Boitani 2003, pp. 261–263.
^ Mech & Boitani 2003, pp. 263–264.
^ Mech & Boitani 2003, p. 266.
^ a b Mech & Boitani 2003, p. 265.
^ Sunquist, Melvin E.; Sunquist, Fiona (2002). Wild cats of the world. University of Chicago Press. p. 167. ISBN 0-226-77999-8.
^ Mech & Boitani 2003, pp. 264–265.
^ Jimenez, Michael D.; Asher, Valpa J.; Bergman, Carita; Bangs, Edward E.; Woodruff, Susannah P. (2008). "Gray Wolves, Canis lupus, Killed by Cougars, Puma concolor, and a Grizzly Bear, Ursus arctos, in Montana, Alberta, and Wyoming". The Canadian Field-Naturalist. 122: 76. doi:10.22621/cfn.v122i1.550.
^ Elbroch, L. M.; Lendrum, P. E.; Newsby, J.; Quigley, H.; Thompson, D. J. (2015). "Recolonizing wolves influence the realized niche of resident cougars". Zoological Studies. 54 (41): e41. doi:10.1186/s40555-015-0122-y. PMC 6661435. PMID 31966128.
^ Miquelle, D. G.; Stephens, P. A.; Smirnov, E. N.; Goodrich, J. M.; Zaumyslova, O. J.; Myslenkov, A. E. (2005). "Tigers and Wolves in the Russian Far East: Competitive Exclusion, Functional Redundancy and Conservation Implications". In Ray, J. C.; Berger, J.; Redford, K. H.; Steneck, R. (eds.). Large Carnivores and the Conservation of Biodiversity. Island Press. pp. 179–207. ISBN 1-55963-080-9. Archived from the original on June 3, 2016. Retrieved November 22, 2015.
^ Monchot, H.; Mashkour, H. (2010). "Hyenas around the cities. The case of Kaftarkhoun (Kashan- Iran)". Journal of Taphonomy. 8 (1): 17–32..
^ Mills, M. G. L.; Mills, Gus; Hofer, Heribert (1998). Hyaenas: status survey and conservation action plan. IUCN. pp. 24–25. ISBN 978-2-8317-0442-5. Archived from the original on May 16, 2016. Retrieved November 22, 2015.
![]()
^ Nayak, S.; Shah, S.; Borah, J. (2015). "Going for the kill: an observation of wolf-hyaena interaction in Kailadevi Wildlife Sanctuary, Rajasthan, India". Canid Biology & Conservation. 18 (7): 27–29.
^ Dinets, Vladimir; Eligulashvili, Beniamin (2016). "Striped Hyaenas (Hyaena hyaena) in Grey Wolf (Canis lupus) packs: Cooperation, commensalism or singular aberration?". Zoology in the Middle East. 62: 85–87. doi:10.1080/09397140.2016.1144292. S2CID 85957777.
^ Mech & Boitani 2003, pp. 208–211.
^ a b Mech & Boitani 2003, pp. 211–213.
^ Graves 2007, pp. 77–85.
^ a b Mech & Boitani 2003, pp. 202–208.
^ Mech & Boitani 2003, p. 164.
^ a b Mech & Boitani 2003, pp. 2–3, 28.
^ Molnar, B.; Fattebert, J.; Palme, R.; Ciucci, P.; Betschart, B.; Smith, D. W.; Diehl, P. (2015). "Environmental and intrinsic correlates of stress in free-ranging wolves". PLOS ONE. 10 (9). e0137378. Bibcode:2015PLoSO..1037378M. doi:10.1371/journal.pone.0137378. PMC 4580640. PMID 26398784.
^ Mech & Boitani 2003, pp. 1–2.
^ Mech & Boitani 2003, pp. 12–13.
^ Nowak, R. M.; Paradiso, J. L. (1983). "Carnivora;Canidae". Walker's Mammals of the World. Vol. 2 (4th ed.). Johns Hopkins University Press. p. 953. ISBN 978-0-8018-2525-5.
^ Mech & Boitani 2003, p. 38.
^ a b Jędrzejewski, W. O.; Schmidt, K.; Theuerkauf, J. R.; Jędrzejewska, B. A.; Kowalczyk, R. (2007). "Territory size of wolves Canis lupus: Linking local (Białowieża Primeval Forest, Poland) and Holarctic-scale patterns". Ecography. 30 (1): 66–76. Bibcode:2007Ecogr..30...66J. doi:10.1111/j.0906-7590.2007.04826.x. S2CID 62800394.
^ a b c Mech & Boitani 2003, pp. 19–26.
^ Mech, L. D. (1977). "Wolf-Pack Buffer Zones as Prey Reservoirs". Science. 198 (4314): 320–321. Bibcode:1977Sci...198..320M. doi:10.1126/science.198.4314.320. PMID 17770508. S2CID 22125487. Archived from the original on July 24, 2018. Retrieved January 10, 2019.
^ Mech, L. David; Adams, L. G.; Meier, T. J.; Burch, J. W.; Dale, B. W. (2003). "Ch.8-The Denali Wolf-Prey System". The Wolves of Denali. University of Minnesota Press. p. 163. ISBN 0-8166-2959-5.
^ Mech & Boitani 2003, pp. 66–103.
^ Busch 2007, p. 59.
^ Lopez 1978, p. 38.
^ Lopez 1978, pp. 39–41.
^ Mech & Boitani 2003, p. 90.
![]()
^ Peters, R. P.; Mech, L. D. (1975). "Scent-marking in wolves". American Scientist. 63 (6): 628–637. Bibcode:1975AmSci..63..628P. PMID 1200478.
^ a b c Heptner & Naumov 1998, p. 248.
^ Mech & Boitani 2003, p. 5.
^ Mech & Boitani 2003, p. 175.
^ Heptner & Naumov 1998, pp. 234–238.
^ a b Mech & Boitani 2003, pp. 42–46.
^ a b Heptner & Naumov 1998, pp. 249–254.
^ Mech & Boitani 2003, p. 47.
^ Mech & Boitani 2003, pp. 46–49.
^ Fox, M. W. (1978). The Dog: Its Domestication and Behavior. Garland STPM. p. 33. ISBN 978-0-89464-202-9.
^ Mech & Boitani 2003, pp. 119–121.
^ Thurber, J. M.; Peterson, R. O. (1993). "Effects of Population Density and Pack Size on the Foraging Ecology of Gray Wolves". Journal of Mammalogy. 74 (4): 879–889. doi:10.2307/1382426. JSTOR 1382426. S2CID 52063038.
^ Mech, Smith & MacNulty 2015, p. 4.
^ Heptner & Naumov 1998, p. 233.
^ Heptner & Naumov 1998, p. 240.
^ MacNulty, Daniel; Mech, L. David; Smith, Douglas W. (2007). "A proposed ethogram of large-carnivore predatory behavior, exemplified by the wolf". Journal of Mammalogy. 88 (3): 595–605. Bibcode:2007JMamm..88..595M. doi:10.1644/06-MAMM-A-119R1.1.
^ a b Mech, Smith & MacNulty 2015, pp. 1–3.
^ Mech, Smith & MacNulty 2015, p. 7.
^ Mech, Smith & MacNulty 2015, p. 82–89.
^ Zimen, Erik (1981). The Wolf: His Place in the Natural World. Souvenir Press. pp. 217–218. ISBN 978-0-285-62411-5.
^ Mech 1981, p. 185.
^ Mech & Boitani 2003, p. 58.
![]()
^ Mech & Boitani 2003, pp. 122–125.
^ Mech & Boitani 2003, p. 230.
^ a b Mech & Boitani 2003, pp. 321–324.
^ Government of Canada (July 29, 2019). "Schedule 3 (section 26) Protected Species". Justice Laws Website. Archived from the original on April 9, 2019. Retrieved October 30, 2019.
^ State of Alaska (October 29, 2019). "Wolf Hunting in Alaska". Alaska Department of Fish and Game. Archived from the original on September 30, 2019. Retrieved October 30, 2019.
^ "Wolf Recovery under the Endangered Species Act" (PDF). US Fish and Wildlife Service. February 2007. Archived (PDF) from the original on August 3, 2019. Retrieved September 1, 2019.
^ "Wolf Numbers in Minnesota, Wisconsin and Michigan (excluding Isle Royale)—1976 to 2015". U.S. Fish and Wildlife Service. Retrieved March 23, 2020.
^ "How many wild wolves are in the United States?". Wolf Conservation Center. Retrieved May 10, 2023.
^ Nie, M. A. (2003). Beyond Wolves: The Politics of Wolf Recovery and Management. University of Minnesota Press. pp. 118–119. ISBN 0-8166-3978-7.
^ "Mexican Wolf Population Grows for Eighth Consecutive Year | U.S. Fish & Wildlife Service". www.fws.gov. March 5, 2024. Retrieved March 6, 2024.
^ Blanco, J.; Sundseth, K. (2023). The situation of the wolf (canis lupus) in the European Union – An in-depth analysis. Publications Office of the European Union. doi:10.2779/187513. ISBN 978-92-68-10281-7. Retrieved February 7, 2025.
^ European Commission: Status, management and distribution of large carnivores—bear, lynx, wolf & wolverine—in Europe December 2012. page 50.
^ Mech & Boitani 2003, pp. 318–320.
^ "Status of large carnivore populations in Europe 2012–2016". European Commission. Archived from the original on September 2, 2019. Retrieved September 2, 2019.
^ The Wolf Dilemma: Following the Practices of Several Actors in Swedish Large Carnivore Management. Ramsey, Morag (2015) https://www.diva-portal.org/smash/get/diva2:821120/FULLTEXT01.pdf Retrieved 30 September 2023
^ Goldthorpe, Gareth (2016). The wolf in Eurasia—a regional approach to the conservation and management of a top-predator in Central Asia and the South Caucasus. Fauna & Flora International. doi:10.13140/RG.2.2.10128.20480.
^ Baskin, Leonid (2016). "Hunting as Sustainable Wildlife Management". Mammal Study. 41 (4): 173–180. doi:10.3106/041.041.0402.
^ "The Wolf in Russia—situations and problems" (PDF). Wolves and Humans Foundation. Archived from the original (PDF) on September 23, 2007. Retrieved September 2, 2019.
^ a b Fisher, A. (January 29, 2019). "Conservation in conflict: Advancement and the Arabian wolf". Middle East Eye. Archived from the original on November 7, 2019. Retrieved November 11, 2019.
^ Mech & Boitani 2003, pp. 323, 327.
^ Busch 2007, p. 231.
^ Şekercioğlu, Çağan (December 15, 2013). "Turkey's Wolves Are Texting Their Travels to Scientists". National Geographic. Archived from the original on October 6, 2019. Retrieved November 19, 2019.
![]()
^ a b Kabir, Muhammad; Hameed, Shoaib; Ali, Hussain; Bosso, Luciano; Din, Jaffar Ud; Bischof, Richard; Redpath, Steve; Nawaz, Muhammad Ali (November 9, 2017). "Habitat suitability and movement corridors of grey wolf (Canis lupus) in Northern Pakistan". PLOS ONE. 12 (11) e0187027. Bibcode:2017PLoSO..1287027K. doi:10.1371/journal.pone.0187027. ISSN 1932-6203. PMC 5679527. PMID 29121089. This article incorporates text available under the CC BY 4.0 license.
^ Mech & Boitani 2003, p. 327.
^ Yadvendradev, V. Jhala; Giles, Robert H. Jr. (991). "The Status and Conservation of the Wolf in Gujarat and Rajasthan, India". Conservation Biology. 5 (4). Wiley: 476–483. Bibcode:1991ConBi...5..476J. doi:10.1111/j.1523-1739.1991.tb00354.x. JSTOR 2386069.
^ Sharma, Lalit Kumar; Mukherjee, Tanoy; Saren, Phakir Chandra; Chandra, Kailash (2019). "Identifying suitable habitat and corridors for Indian Grey Wolf (Canis lupus pallipes) in Chotta Nagpur Plateau and Lower Gangetic Planes: A species with differential management needs". PLOS ONE. 14 (4) e0215019. Bibcode:2019PLoSO..1415019S. doi:10.1371/journal.pone.0215019. PMC 6457547. PMID 30969994.
^ Mech & Boitani 2003, p. 328.
^ Larson, Greger (2017). "Reconsidering the distribution of gray wolves". Zoological Research. 38 (3): 115–116. doi:10.24272/j.issn.2095-8137.2017.021. PMC 5460078. PMID 28585433.
^ Busch 2007, p. 232.
^ Xu, Yu; Yang, Biao; Dou, Liang (2015). "Local villagers' perceptions of wolves in Jiuzhaigou County, western China". PeerJ. 3 e982. doi:10.7717/peerj.982. PMC 4465947. PMID 26082870.
^ Ishiguro, Naotaka; Inoshima, Yasuo; Shigehara, Nobuo (2009). "Mitochondrial DNA Analysis of the Japanese Wolf (Canis Lupus Hodophilax Temminck, 1839) and Comparison with Representative Wolf and Domestic Dog Haplotypes". Zoological Science. 26 (11): 765–70. doi:10.2108/zsj.26.765. PMID 19877836. S2CID 27005517.
^ a b Mech & Boitani 2003, p. 292.
^ Lopez 1978, p. 210.
^ Marvin 2012, p. 128.
^ a b c Werness, Hope B. (2007). The Continuum Encyclopedia of Animal Symbolism in World Art. Continuum International Publishing Group. pp. 405, 437. ISBN 978-0-8264-1913-2.
^ Marvin 2012, p. 78.
^ Lopez 1978, p. 133.
^ Busch 2007, p. 110.
^ Marvin 2012, p. 47.
^ Marvin 2012, p. 50.
^ Lopez 1978, p. 123.
^ Marvin 2012, pp. 38–45.
^ Lopez 1978, p. 259.
^ Marvin 2012, pp. 64–70.
![]()
^ Lopez 1978, p. 263.
^ Graves 2007, pp. 21, 123.
^ Marvin 2012, p. 162.
^ Cassidy, K; Smith, D. W.; Mech, L. D.; MacNulty, D. R.; Stahler, D. R.; Metz, M. C. (2006). "Territoriality and interpack aggression in wolves: Shaping a social carnivore's life history. Rudyard Kipling's Law of the Jungle Meets Yellowstone's Law of the Mountains". Yellowstone Science. 24 (1): 37–41.
^ Mech & Boitani 2003, p. 294.
^ Jones, K. (2001). "Never Cry Wolf: Science, Sentiment, and the Literary Rehabilitation of Canis Lupus". The Canadian Historical Review. 84. Archived from the original on October 12, 2013. Retrieved July 28, 2012.
^ Grooms, Steve (2008). "The Mixed Legacy of Never Cry Wolf" (PDF). International Wolf. 18 (3): 11–13. Archived from the original (PDF) on June 21, 2010.
^ Creel, S.; Fox, J. E.; Hardy, A.; Sands, J.; Garrott, B.; Peterson, R. O. (2002). "Snowmobile activity and glucocorticoid stress responses in wolves and elk". Conservation Biology. 16 (3): 809–814. Bibcode:2002ConBi..16..809C. doi:10.1046/j.1523-1739.2002.00554.x. S2CID 84878446.
^ a b c d Mech & Boitani 2003, p. 305.
^ Mech & Boitani 2003, p. 309.
^ Mech 1981, p. 173.
^ Levy, Max G. (December 11, 2020). "These Non-Lethal Methods Encouraged by Science Can Keep Wolves From Killing Livestock". Smithsonian Magazine. Retrieved December 16, 2020.
^ Mech & Boitani 2003, p. 307.
^ Mech & Boitani 2003, p. 306.
^ Graves 2007, p. 45.
^ Vanak, A. T.; Dickman, C. R.; Silva-Rodriguez, E. A.; Butler, J. R. A.; Ritchie, E. G. (2014). "Top-dogs and under-dogs: competition between dogs and sympatric carnivores". In Gompper, M. E. (ed.). Free-Ranging Dogs and Wildlife Conservation. Oxford University Press. pp. 69–93. ISBN 978-0-19-966321-7.
^ a b c d e f g h Lescureux, Nicolas; Linnell, John D. C. (2014). "Warring brothers: The complex interactions between wolves (Canis lupus) and dogs (Canis familiaris) in a conservation context". Biological Conservation. 171: 232–245. Bibcode:2014BCons.171..232L. doi:10.1016/j.biocon.2014.01.032.
^ Boitani, L. (1983). "Wolf and dog competition in Italy". Acta Zoologica Fennica (174): 259–264.
^ Backeryd, J. (2007). "Wolf attacks on dogs in Scandinavia 1995–2005—Will wolves in Scandinavia go extinct if dog owners are allowed to kill a wolf attacking a dog?" (PDF). Examensarbete, Institutionen för ekologi, Grimsö forskningsstation. Sveriges Lantbruksuniversitet. Archived from the original (PDF) on July 17, 2019. Retrieved July 17, 2019.
^ Shivik, John A. (2006). "Tools for the Edge: What's New for Conserving Carnivores". BioScience. 56 (3): 253. doi:10.1641/0006-3568(2006)056[0253:TFTEWN]2.0.CO;2.
^ Coppinger, R.; Schneider, R. (1995). "Evolution of working dogs". In Serpell, J. (ed.). The Domestic Dog: Its Evolution, Behaviour and Interactions With People. University Press, Cambridge. pp. 21–47. ISBN 978-0-521-42537-7.
^ Espuno, Nathalie; Lequette, Benoit; Poulle, Marie-Lazarine; Migot, Pierre; Lebreton, Jean-Dominique (2004). "Heterogeneous response to preventive sheep husbandry during wolf recolonization of the French Alps". Wildlife Society Bulletin. 32 (4): 1195–1208. doi:10.2193/0091-7648(2004)032[1195:HRTPSH]2.0.CO;2. S2CID 86058778.
^ a b c d e f Linnell, J. D. C. (2002). The Fear of Wolves: A Review of Wolf Attacks on Humans (PDF). Norsk Institutt for Naturforskning (NINA). ISBN 978-82-426-1292-2. Archived (PDF) from the original on May 17, 2014. Retrieved August 16, 2013.
^ Mech & Boitani 2003, pp. 300–304.
^ Rajpurohit, K.S. (1999). "Child lifting: Wolves in Hazaribagh, India". Ambio. 28: 162–166.
^ Heptner & Naumov 1998, p. 267.
^ Roosevelt, Theodore (1909). Hunting the grisly and other sketches; an account of the big game of the United States and its chase with horse, hound, and rifle. G. P. Putnam's sons. pp. 179–207. Archived from the original on June 24, 2015. Retrieved May 14, 2014.
^ Lopez 1978, p. 108.
^ a b Graves 2007, pp. 121–140.
^ Tucker, P.; Weide, B. (1998). "Can You Turn a Wolf into a Dog" (PDF). Wild Sentry. Archived from the original (PDF) on December 8, 2015. Retrieved March 21, 2016.
Busch, R. H. (2007). Wolf Almanac, New and Revised: A Celebration Of Wolves And Their World (3 ed.). Rowman & Littlefield. ISBN 978-1-59921-069-8.
Graves, Will (2007). Wolves in Russia: Anxiety throughout the ages. Detselig Enterprises. ISBN 978-1-55059-332-7.
Heptner, V. G.; Naumov, N. P. (1998). Mammals of the Soviet Union Vol. II Part 1a, Sirenia and Carnivora (Sea cows; Wolves and Bears). Science Publishers, Inc. USA. ISBN 978-1-886106-81-9.
Lopez, Barry H. (1978). Of Wolves and Men. J. M. Dent and Sons Limited. ISBN 978-0-7432-4936-2.
Marvin, Garry (2012). Wolf. Reaktion Books Ltd. ISBN 978-1-86189-879-1.
Mech, L. David (1981). The Wolf: The Ecology and Behaviour of an Endangered Species. University of Minnesota Press. ISBN 978-0-8166-1026-6.
Mech, L. David; Boitani, Luigi, eds. (2003). Wolves: Behaviour, Ecology and Conservation. University of Chicago Press. ISBN 978-0-226-51696-7.
Mech, L. David; Smith, Douglas W.; MacNulty, Daniel R. (2015). Wolves on the Hunt: The Behavior of Wolves Hunting Wild Prey. University of Chicago Press. ISBN 978-0-226-25514-9.
Wikiquote has quotations related to Wolf.
The International Wolf Center
Staying Safe in Wolf Country, ADFG (January 2009)
Watch Death of a Legend and Cry of the Wild Archived April 20, 2009, at the Wayback Machine by Bill Mason
🔐 Cryptographic Verification
Archived URL: https://en.wikipedia.org/wiki/Wolf
📄 CONTENT HASHES:
SHA-256: 46b03a813204e2281a93bde7beb900be6db17cbd560fc481387c5f2f57e1b819
BLAKE2b: 4d8b00c6dce171ff80d7ac4b1c518704258b97dc146f3d50589135a3f103b929
MD5: e7b26d3b280003a9e540a62326d633c7
📝 TITLE HASHES:
SHA-256: ea858afbf837aae714617e89a36524aced28f7de921f7798e72810fd8839a462
BLAKE2b: 074a64d52218ca49adc8c8ca4abc182cd4fdbeadb47d9f02f1bcb14d9c3b3d78
MD5: 279ede0c6bdbd7d26309becd159053d8
🔒 INTEGRITY HASHES:
SHA-256: 27751fe0f2b84333b9f142a3ba68932bf675c4012ca11cac3a5b607c6499ce58
BLAKE2b: d1d3b7fa1f0610a3db1e460dc32b845abae833f5166d8661fbb7bef7c240b3dd
MD5: 552618e783978ef57d2c765812601c82
(html comment removed: ARCHIVE-MANIFEST:{"s":"ebcd14fd-bfc3-46de-819f-56469dd730dc","v":"1.0","t":3,"p":2,"h":{"sha256":"31d87832e9de7fb3f245cff5a48e6cc4d3f68b4a35d5e9d4a52806e1e5f4f508","blake2b":"307ba8842bb064270c54f5fe14aa3c7f475c9a93e5a41abc36cf85a70986d4bcd24fd5dba041a74242a99d594d7ffed05fa34df03ee991e1a6932a339912361a","md5":"1acdf8b4bc0619e0f92ec8b660fac20e"},"u":"https://en.wikipedia.org/wiki/Wolf"})