I looked again at an earlier study by Fliermans & Schmidt (1977) that demonstrates the presence of chemolithotrophic nitrifying bacteria (Nitrobacter sp.) using species-specific fluorescent antibodies to show Nitrobacter cells attached to sediment particles associated with saltpeter mining:
http://scholarcommons.usf.edu/cgi/viewcontent.cgi?article=1493&context=ijs
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